Lizard

Description

Most lizards retain the typical tetrapod body plan of a short neck, four limbs of roughly equal size ending in five toes each, a moderately long body, and a long tail. Most lizards possess external ears and have movable eyelids. Encompassing forty families, there is tremendous variety in color, appearance and size. Many are also capable of regenerating tails. Almost all lizards are carnivorous, though most are so small that insects are their primary prey. About 120 species, mostly iguanids, are known to be herbivorous. A few species are omnivorous, and others have reached sizes where they can prey on other vertebrates. Many lizards are good climbers or fast sprinters. Some can run bipedally, such as the collared lizard, and some can even run across the surface of water to escape, notably the basilisk. The fastest lizard is the Costa Rican spiny-tailed iguana (Ctenosaura similis), which can sprint at up to 21.5 miles/h (34.6 km/h).[1]

Many lizards can change colour in response to their environments or in times of stress. The most familiar example is the chameleon, but more subtle colour changes occur in other lizard species as well, such as the anole, also known as the "American chameleon", "house chameleon" or "chameleon."

Lizards have relatively low rates of metabolism, as compared with birds or mammals, and are termed ectothermic, meaning that they rely on external sources for heat. Many species bask in the sun to raise their body temperature. By this behavioral thermoregulation mechanism, many species are able to maintain high and quite stable body temperatures while they are active. At night, however, or during cold times of the year, behavioral thermoregulation cannot achieve high body temperatures, so their temperature tends to vary with that of their surroundings, and lizards are also termed poikilothermic. Lizards are most diverse and abundant in areas with consistently high temperatures, including many deserts. Lizards are rarely seen in the upper half of the United States and most European countries.

[edit] Senses and communication

Lizards employ many diverse methods of communication. Like many other animals, they have an acute sense of smell, detecting scents of their prey or pheromones from other lizards. The primary organ of scent in lizards is a vomeronasal organ in the roof of the mouth, and lizards gather scents by flicking out their tongues, then retracting them and delivering the captured odor molecules to this organ. Some large carnivorous lizards, such as tegus and monitor lizards, have forked tongues like snakes, to take advantage of this organ better. As a result, many male lizards possess enlarged pores on the underside of their thighs, which they rub against objects to mark their territory.

While most lizards can hear well, few are capable of vocalizations or otherwise making noise. The exception to this rule is the geckos, which communicate through a wide variety of barks, chirps and whistles, with each species having specific patterns and sounds.

Sight is quite important for most lizards, both for locating prey and for communication, and as such, many lizards have highly acute color vision. Most lizards rely heavily on body language, using specific postures, gestures and movements to define territory, resolve disputes, and entice mates. Some species of lizard also utilize bright colors, such as the iridescent patches on the belly of Sceloporus. These colors would be highly visible to predators, so are often hidden on the underside or between scales and only revealed when necessary.

A particular innovation in this respect is the dewlap, a brightly colored patch of skin on the throat, usually hidden between scales. When a display is needed, the lizards erect the hyoid bone of their throat, resulting in a large vertical flap of brightly colored skin beneath the head which can be then used for communication. Anoles are particularly famous for this display, with each species having specific colors, including patterns only visible under ultraviolet light, as lizards can often see UV.

[edit] Evolution and relationships
Frilly-necked lizard Chlamydosaurus kingii

The retention of the basic tetrapod body form by lizards makes it tempting to assume any similar animal, alive or extinct, is also a lizard. However, this is not the case, and lizards are part of a well-defined group.

The first reptile was superficially lizard-like, but had a solid, box-like skull, with openings only for eyes, nostrils, etc (termed Anapsid). These organisms later gave rise to two new groups with additional holes in the skull to make room for and anchor larger jaw muscles.Those with a single hole, the Synapsids, became modern mammals. The Diapsids, possessing two holes, continued to diversify. The Archosaurs retained the basic Diapsid skull, and gave rise to a bewildering array of animals, most famous being the dinosaurs and their descendants, birds. The Lepidosaurs began to reduce the skull bones, making the skull lighter and more flexible. Modern tuataras retain the basic Lepidosaur skull, distinguishing them from true lizards in spite of superficial similarities. Squamates, including snakes and all true lizards, further lightened the skull by eliminating the lower margin of the lower skull opening.